Y. Han et al.
Citric acid
Acetic acid
Aspartate
Acetic acid
Succinic acid
Ethanol
Higher alcohols
Malate
→Malic acid
Fumaric acid
Il
-Succinic acid
Diacetyl
Diacetyl
a-acetolactate
Amino acid
Keto acids
Ethanol
Higher alcohols
Acetaldehyde
TCA
Pyravate carbylyTuvic acid-
Oxaloacetate Acetyl-CoA
ketoglutarate
Sugar
Acetaldehyde
+
1
I
1
T
T
complea-ketoglutarate
Fig. 1. The main metabolism of organic acids by yeast.
Esters
Esters
EMP
Isocitrate-
Citric acid Glyoxylate cycle Malic acid
Glutamate
Sugar
EMP
L--Acetaldehyde-
Acetyl COA
Fatty acid CoA.
Fatty Acids
Oxaloacetate
Fatty Acids
Lactic acid
Lactic acid
Pyruvic acid-Lactic acid Ethyl lactate
Ethyl lactate
Ethanol-
Glyoxylate
Succinic acid.
Fig. 2. The main metabolism of volatiles by yeast.
Ethyl lactate
Acetic acid
Succinic acid
Ethyl esters-
Acetic acid
Ethylacetate
Malic acid
Ethyl esters
LWT 173 (2023) 114389